T.  Le Péchon, Dubuisson, J. - Y., Haevermans, T., Cruaud, C., Couloux, A., et Gigord, L. D. B., « Multiple colonizations from Madagascar and converged acquisition of dioecy in the Mascarene Dombeyoideae (Malvaceae) as inferred from chloroplast and nuclear DNA sequence analyses. », Annals of botany, vol. 106, p. 343?57, 2010.